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Field Descriptions
228 matching records found.
Label Query: Genus = "Dryopteris", Species = "expansa"
Dryopteridaceae
1. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Russian Federation, Sakhalin Region
Sakhalin Island, on Schmidta Peninsula at northern end of island; along Sukharnyy River ca 7 km SSW of Tri Brata Mountain.
Elev. 394 ft. 54° 2' 2.58" N, 142° 48' 21" E.
Dense Picea-Betula forest, with understory of Cornus canadensis, ferns, mosses; also some small logging clearings and buildings. Fronds 1-3 per rhizome; abundant in forest. Phenology: Fertile. Origin: Native.
WTU 357906
Ben Legler 1176    9 Aug 2003
1. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
872 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
357906 
Barcode
 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Slava Barkalov, 2003.  
Type Notes
 
Collector
Ben Legler 
Collection #
1176 
Other Collectors
 
Date Collected
9 Aug 2003 
Verbatim Date
 
Country
Russian Federation 
State or Province
Sakhalin Region 
County
 
Locality
Sakhalin Island, on Schmidta Peninsula at northern end of island; along Sukharnyy River ca 7 km SSW of Tri Brata Mountain. 
Elevation
394 ft. 
Coordinates
54° 2' 2.58" N, 142° 48' 21" E 
Site Description
Dense Picea-Betula forest, with understory of Cornus canadensis, ferns, mosses; also some small logging clearings and buildings. 
Specimen Notes
Fronds 1-3 per rhizome; abundant in forest. 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Wallowa County:
Wallowa-Whitman National Forest. Lostine Canyon, Bowman Trail, where trail first crosses a feeder creek.
Elev. 6689 ft. 45° 17.289' N, 117° 24.645' W.
Dry, north-facing outcrop; open area, break in trees. Granite cliff and talus. Scales on petiole rust-red. Altimeter reading 6450 ft.. Phenology: Vegetative. Origin: Native.
WTU 360641; Barcode: WTU-V-015504
Sarah Gage 7347    21 Jul 2002
with Don Knoke, Dale Blum, Richard Robohm
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
1414 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
360641 
Barcode
WTU-V-015504 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ben Legler, 19 2002. Sources: H&C.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 19 2008. Sources: FNA.  
Type Notes
 
Collector
Sarah Gage 
Collection #
7347 
Other Collectors
Don Knoke, Dale Blum, Richard Robohm 
Date Collected
21 Jul 2002 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Wallowa 
Locality
Wallowa-Whitman National Forest. Lostine Canyon, Bowman Trail, where trail first crosses a feeder creek. 
Elevation
6689 ft. 
Coordinates
45° 17.289' N, 117° 24.645' W 
Site Description
Dry, north-facing outcrop; open area, break in trees. Granite cliff and talus. 
Specimen Notes
Scales on petiole rust-red. Altimeter reading 6450 ft.. 
Phenology
Vegetative 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Island County:
Whidbey Island, Ebey's Landing; Hill Road c. 1 km South of main parking lot, just below summit of road. Bluff to west of road.
Elev. 171 ft. 48° 11' 17" N, 122° 42' 5" W. Datum: NAD 27.
Steep bluff dominated by native and exotic grasses with Lupinus, Eriophyllum, and Achillea; backed by conifer woods. In woods behind bluff. SHEET 1 of 2. Phenology: Fruits. Origin: Native.
WTU 358070; Barcode: WTU-V-015413
Peter W. Dunwiddie PWD 78    30 Apr 2004
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
1504 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
358070 
Barcode
WTU-V-015413 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ben Legler, 20 2004. Sources: FNA.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU. 
Type Notes
 
Collector
Peter W. Dunwiddie 
Collection #
PWD 78 
Other Collectors
 
Date Collected
30 Apr 2004 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Island 
Locality
Whidbey Island, Ebey's Landing; Hill Road c. 1 km South of main parking lot, just below summit of road. Bluff to west of road. 
Elevation
171 ft. 
Coordinates
48° 11' 17" N, 122° 42' 5" W 
Site Description
Steep bluff dominated by native and exotic grasses with Lupinus, Eriophyllum, and Achillea; backed by conifer woods. 
Specimen Notes
In woods behind bluff. SHEET 1 of 2. 
Phenology
Fruits 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
4. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Island County:
Whidbey Island, Ebey's Landing; Hill Road c. 1 km South of main parking lot, just below summit of road. Bluff to west of road.
Elev. 171 ft. 48° 11' 17" N, 122° 42' 5" W. Datum: NAD 27.
Steep bluff dominated by native and exotic grasses with Lupinus, Eriophyllum, and Achillea; backed by conifer woods. In woods behind bluff. SHEET 2 of 2. Phenology: Fruits. Origin: Native.
WTU 358071; Barcode: WTU-V-015412
Peter W. Dunwiddie PWD 78    30 Apr 2004
4. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
1593 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
358071 
Barcode
WTU-V-015412 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ben Legler, 20 2004. Sources: FNA.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
Peter W. Dunwiddie 
Collection #
PWD 78 
Other Collectors
 
Date Collected
30 Apr 2004 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Island 
Locality
Whidbey Island, Ebey's Landing; Hill Road c. 1 km South of main parking lot, just below summit of road. Bluff to west of road. 
Elevation
171 ft. 
Coordinates
48° 11' 17" N, 122° 42' 5" W 
Site Description
Steep bluff dominated by native and exotic grasses with Lupinus, Eriophyllum, and Achillea; backed by conifer woods. 
Specimen Notes
In woods behind bluff. SHEET 2 of 2. 
Phenology
Fruits 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
5. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Skamania County:
Refugia area between Pumice Plain and Plains of Abraham, northeast of crater breach. Near Spirit Lake.
46.27333°, -122.13389°. Datum: WGS 84. Coordinate uncertainty: 5000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Origin: Native.
WTU 334498; Barcode: WTU-V-015617
Jon Titus s.n.    15 Jul 1994
5. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
6784 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
334498 
Barcode
WTU-V-015617 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Athyrium filix-femina (L.) Roth.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
Jon Titus 
Collection #
s.n. 
Other Collectors
 
Date Collected
15 Jul 1994 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Skamania 
Locality
Refugia area between Pumice Plain and Plains of Abraham, northeast of crater breach. Near Spirit Lake. 
Elevation
 
Coordinates
46.27333°, -122.13389° 
Site Description
 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
6. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Thurston County:
Priest Point Park, Olympia. East of East Bay Drive, southern part of park.
Elev. 100-120 ft. 47° 4' N, 122° 53' W; T18N R2W S2 or 11.
Origin: Native.
WTU 345073; Barcode: WTU-V-015616
Sarah Gage G&R 037    22 May 1992
with Sharon Rodman
6. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
6785 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
345073 
Barcode
WTU-V-015616 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Athyrium filix-femina (L.) Roth.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
Sarah Gage 
Collection #
G&R 037 
Other Collectors
Sharon Rodman 
Date Collected
22 May 1992 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Thurston 
Locality
Priest Point Park, Olympia. East of East Bay Drive, southern part of park. 
Elevation
100-120 ft. 
Coordinates
47° 4' N, 122° 53' W; T18N R2W S2 or 11 
Site Description
 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
7. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Lewis County:
Ohanapecosh Hot Springs, Mount Rainier National Park.
Elev. 3000 ft. 46.736944°, -121.559167°. Datum: NAD 27. Coordinate uncertainty: 200 meters. Georeferenced by WTU Staff. Coordinate Source: Georeferenced.
Deep woods in moist places. Phenology: Spores. Origin: Native.
WTU 204JWT; Barcode: WTU-V-015648
J. William Thompson 12680    27 Aug 1935
7. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
6832 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
204JWT 
Barcode
WTU-V-015648 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946. Remarks: Herbarium of the University of California.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
J. William Thompson 
Collection #
12680 
Other Collectors
 
Date Collected
27 Aug 1935 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Lewis 
Locality
Ohanapecosh Hot Springs, Mount Rainier National Park. 
Elevation
3000 ft. 
Coordinates
46.736944°, -121.559167° 
Site Description
 
Specimen Notes
Deep woods in moist places. 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
8. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Jefferson County:
Along Quinault River, about 15 miles north of south boundary of Olympic National Park.
47.57033°, -123.58617°. Datum: WGS 84. Coordinate uncertainty: 2000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Phenology: Spores. Origin: Native.
WTU 97232; Barcode: WTU-V-015652
H. T. Rogers 817    23 Jun 1941
8. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
6834 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
97232 
Barcode
WTU-V-015652 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
H. T. Rogers 
Collection #
817 
Other Collectors
 
Date Collected
23 Jun 1941 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Jefferson 
Locality
Along Quinault River, about 15 miles north of south boundary of Olympic National Park. 
Elevation
 
Coordinates
47.57033°, -123.58617° 
Site Description
 
Specimen Notes
 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
9. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Mason County:
1/2 mile east of Satsop-Shelton Rd., opposite N end of Schafer Park.
47.215272°, -123.056817°. Datum: WGS 84. Coordinate uncertainty: 5000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Growing in rotten fir stump. Phenology: Spores. Origin: Native.
WTU 127108; Barcode: WTU-V-015649
Phillips B. Freer 77    12 Aug 1949
9. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
6836 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
127108 
Barcode
WTU-V-015649 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
Phillips B. Freer 
Collection #
77 
Other Collectors
 
Date Collected
12 Aug 1949 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Mason 
Locality
1/2 mile east of Satsop-Shelton Rd., opposite N end of Schafer Park. 
Elevation
 
Coordinates
47.215272°, -123.056817° 
Site Description
 
Specimen Notes
Growing in rotten fir stump. 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
10. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Lewis County:
White Pass Highway, just east of Packwood.
46.60667°, -121.66944°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Second growth coniferous woods. Common in deep woods. Phenology: Spores. Origin: Native.
WTU 213539; Barcode: WTU-V-015650
W. H. Wagner 63107    26 Jun 1963
with A. R. Kruckeberg
10. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
6837 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
213539 
Barcode
WTU-V-015650 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
W. H. Wagner 
Collection #
63107 
Other Collectors
A. R. Kruckeberg 
Date Collected
26 Jun 1963 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Lewis 
Locality
White Pass Highway, just east of Packwood. 
Elevation
 
Coordinates
46.60667°, -121.66944° 
Site Description
Second growth coniferous woods. 
Specimen Notes
Common in deep woods. 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
11. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Island County:
Cranberry Lake, Deception Pass State Park, Whidbey Island.
48.14389°, -122.49111°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Forest. Phenology: Spores. Origin: Native.
WTU 221426; Barcode: WTU-V-015653
Harold W. Smith 805    3 Jun 1936
11. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
6839 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
221426 
Barcode
WTU-V-015653 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
Harold W. Smith 
Collection #
805 
Other Collectors
 
Date Collected
3 Jun 1936 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Island 
Locality
Cranberry Lake, Deception Pass State Park, Whidbey Island. 
Elevation
 
Coordinates
48.14389°, -122.49111° 
Site Description
Forest. 
Specimen Notes
 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
12. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Pacific County:
Fort Columbia.
46.25308°, -123.92065°. Datum: WGS 84. Coordinate uncertainty: 500 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Damp forest. Phenology: Spores. Origin: Native.
WTU 272662; Barcode: WTU-V-015647
Wilhelm N. Suksdorf 6741    22 Jul 1909
12. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
6840 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
272662 
Barcode
WTU-V-015647 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Aspidium dilatatum (Hoffm.) Sm.
2. Dryopteris austriaca (Jacq.) Woynar.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
Wilhelm N. Suksdorf 
Collection #
6741 
Other Collectors
 
Date Collected
22 Jul 1909 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Pacific 
Locality
Fort Columbia. 
Elevation
 
Coordinates
46.25308°, -123.92065° 
Site Description
Damp forest. 
Specimen Notes
 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
13. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Kitsap County:
Near Mountaineers' Cabin.
47.59746°, -122.73449°. Datum: WGS 84. Coordinate uncertainty: 500 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Rich moist woods near stream. Fronds gracefully curving, about 1 meter high. Phenology: Spores. Origin: Native.
WTU 316086; Barcode: WTU-V-015651
George Neville Jones 4327     Aug 1932
13. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
6842 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
316086 
Barcode
WTU-V-015651 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Athyrium filix-femina (L.) Roth.
    Annotated by G. N. Jones.
2. Dryopteris dilatata auct. non (Hoffm.n) Gray.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
George Neville Jones 
Collection #
4327 
Other Collectors
 
Date Collected
Aug 1932 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Kitsap 
Locality
Near Mountaineers' Cabin. 
Elevation
 
Coordinates
47.59746°, -122.73449° 
Site Description
Rich moist woods near stream. 
Specimen Notes
Fronds gracefully curving, about 1 meter high. 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
14. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Whatcom County:
Hozomeen Ranger Station, ca. 1.0 mi. south. Hozomeen Lake Trail.
Elev. 2025-2050 ft. T40N R13E S1. Datum: NAD 27. Coordinate uncertainty: 500 meters. Georeferenced by WTU Staff. Coordinate Source: Georeferenced.
Scattered on damp shady slope in THPL woods, growing with Oplopanax horridum, mosses. Several fronds in a vaselike clump. Phenology: Spores. Origin: Native.
WTU 316967; Barcode: WTU-V-015646
Ralph Naas 5341    21 Jul 1987
with Dorothy Naas
14. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
6844 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
316967 
Barcode
WTU-V-015646 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Joe Arnett, 7 1987.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
Ralph Naas 
Collection #
5341 
Other Collectors
Dorothy Naas 
Date Collected
21 Jul 1987 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Whatcom 
Locality
Hozomeen Ranger Station, ca. 1.0 mi. south. Hozomeen Lake Trail. 
Elevation
2025-2050 ft. 
Coordinates
T40N R13E S1 
Site Description
Scattered on damp shady slope in THPL woods, growing with Oplopanax horridum, mosses. 
Specimen Notes
Several fronds in a vaselike clump. 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
15. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Clark County:
Along Lacamas Creek, Green Mountain Resort Conservation Easement.
T2N R3E S20, 21 Willamette Meridian. Georeferenced by Ed Alverson. Coordinate Source: Inferred from duplicate specimen.
With Montia perfoliata. Origin: Native.
WTU 339130; Barcode: WTU-V-015645
Eliza Habegger 355    1 May 1998
15. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
6846 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
339130 
Barcode
WTU-V-015645 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
Eliza Habegger 
Collection #
355 
Other Collectors
 
Date Collected
1 May 1998 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Clark 
Locality
Along Lacamas Creek, Green Mountain Resort Conservation Easement. 
Elevation
 
Coordinates
T2N R3E S20, 21 Willamette Meridian 
Site Description
 
Specimen Notes
With Montia perfoliata. 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
16. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Snohomish County:
Heather Lake ca 7 miles east of Granite Falls on northeast side of Mt. Pilchuck.
T30N R8E S21 SE. Datum: NAD 27. Coordinate uncertainty: 805 meters. Georeferenced by Ben Legler. Coordinate Source: TRS2LL.
North facing boulder slope above south end of lake. Phenology: Spores. Origin: Native.
WTU 333216; Barcode: WTU-V-015414
M. Arnot s.n.    1 Aug 1994
16. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
6848 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
333216 
Barcode
WTU-V-015414 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by M. Arnot.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
M. Arnot 
Collection #
s.n. 
Other Collectors
 
Date Collected
1 Aug 1994 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Snohomish 
Locality
Heather Lake ca 7 miles east of Granite Falls on northeast side of Mt. Pilchuck. 
Elevation
 
Coordinates
T30N R8E S21 SE 
Site Description
North facing boulder slope above south end of lake. 
Specimen Notes
 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
17. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, King County:
Cascade Mountains of Western Washington: Cedar River Watershed. Rock Creek wetland system. Head into south end from rd 40. Drop into system at Binker culvert and head into west side of the wetland.
Elev. 800 ft. T22N R7E S16 NE. Datum: NAD 27. Coordinate uncertainty: 500 meters. Georeferenced by WTU Staff. Coordinate Source: Georeferenced.
Phenology: Spores. Origin: Native.
WTU 350630; Barcode: WTU-V-015496
Clayton J. Antieau 01-22    26 May 2001
with Linda Cooley, Kristie Dinkin
17. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
23317 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
350630 
Barcode
WTU-V-015496 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Jennifer French, 23 2002.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Clayton J. Antieau 
Collection #
01-22 
Other Collectors
Linda Cooley, Kristie Dinkin 
Date Collected
26 May 2001 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
King 
Locality
Cascade Mountains of Western Washington: Cedar River Watershed. Rock Creek wetland system. Head into south end from rd 40. Drop into system at Binker culvert and head into west side of the wetland. 
Elevation
800 ft. 
Coordinates
T22N R7E S16 NE 
Site Description
 
Specimen Notes
 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
18. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, King County:
Cascade Mountains of Western Washington: Cedar River bridge at Barneston.
Elev. 1000 ft. 47° 23.33' 33" N, 121° 51.68' 51" W; T22N R7E S14.
Woods, riparian area, and roadside near bridge. In shade near river. Phenology: Fertile. Origin: Native.
WTU 350669; Barcode: WTU-V-015497
P. F. Zika 16039    26 May 2001
with Jeff Walker, David Kiefer, Kari Stices, Bill Winije
18. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
23584 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
350669 
Barcode
WTU-V-015497 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Jennifer French, 23 2002.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
P. F. Zika 
Collection #
16039 
Other Collectors
Jeff Walker, David Kiefer, Kari Stices, Bill Winije 
Date Collected
26 May 2001 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
King 
Locality
Cascade Mountains of Western Washington: Cedar River bridge at Barneston. 
Elevation
1000 ft. 
Coordinates
47° 23.33' 33" N, 121° 51.68' 51" W; T22N R7E S14 
Site Description
Woods, riparian area, and roadside near bridge. 
Specimen Notes
In shade near river. 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
19. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, King County:
Cascade Mountains of Western Washington: Wetlands along road 30; near headwaters of Williams Creek.
Elev. 1000 ft. T22N R7E S2 NE 1/4. Datum: NAD 27. Coordinate uncertainty: 805 meters. Georeferenced by Ben Legler. Coordinate Source: TRS2LL.
Other plants in vicinity: Thuja plicata, Tsuga heterophylla, Psuedotsuga menziesii, Acer circinatum, Galium sp., Glycyrrhiza sp. Phenology: Fertile. Origin: Native.
WTU 350951; Barcode: WTU-V-015498
Alan C. Yen 01-46    26 May 2001
19. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
23607 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
350951 
Barcode
WTU-V-015498 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Jennifer French, 23 2002.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Alan C. Yen 
Collection #
01-46 
Other Collectors
 
Date Collected
26 May 2001 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
King 
Locality
Cascade Mountains of Western Washington: Wetlands along road 30; near headwaters of Williams Creek. 
Elevation
1000 ft. 
Coordinates
T22N R7E S2 NE 1/4 
Site Description
Other plants in vicinity: Thuja plicata, Tsuga heterophylla, Psuedotsuga menziesii, Acer circinatum, Galium sp., Glycyrrhiza sp. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
20. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, King County:
Cascade Mountains of Western Washington: Cedar River.
Elev. 800 ft. 47° 23' 20.81" N, 121° 52' 57.43" W; T22N R7E S14.
Along road 10.3, near powerline crossing, north of the river. Field notes include an extensive plant list. Field notes include an extensive plant list for the area. Phenology: Fertile. Origin: Native.
WTU 351447; Barcode: WTU-V-015500
Richard G. Olmstead 01-26    26 May 2001
with Jeremiah Ralston, Rich Gwodz, Liz Stenning, Mark Joselyn
20. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
24367 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
351447 
Barcode
WTU-V-015500 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Jennifer French, 23 2002.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Richard G. Olmstead 
Collection #
01-26 
Other Collectors
Jeremiah Ralston, Rich Gwodz, Liz Stenning, Mark Joselyn 
Date Collected
26 May 2001 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
King 
Locality
Cascade Mountains of Western Washington: Cedar River. 
Elevation
800 ft. 
Coordinates
47° 23' 20.81" N, 121° 52' 57.43" W; T22N R7E S14 
Site Description
Along road 10.3, near powerline crossing, north of the river. Field notes include an extensive plant list. 
Specimen Notes
Field notes include an extensive plant list for the area. 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
21. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, King County:
Cascade Mountains of Western Washington: Cedar River.
Elev. 601 ft. 47° 23' 28.9" N, 121° 55' 12.47" W; T22N R7E S16 SE 1/4.
Along Cedar River at road 40.3, north of the river. Field notes include an extensive plant list. Phenology: Fertile. Origin: Native.
WTU 351445; Barcode: WTU-V-015499
Richard G. Olmstead 01-12    26 May 2001
with Jeremiah Ralston, Rich Gwodz, Liz Stenning, Mark Joselyn
21. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
24369 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
351445 
Barcode
WTU-V-015499 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Jennifer French, 23 2002.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Richard G. Olmstead 
Collection #
01-12 
Other Collectors
Jeremiah Ralston, Rich Gwodz, Liz Stenning, Mark Joselyn 
Date Collected
26 May 2001 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
King 
Locality
Cascade Mountains of Western Washington: Cedar River. 
Elevation
601 ft. 
Coordinates
47° 23' 28.9" N, 121° 55' 12.47" W; T22N R7E S16 SE 1/4 
Site Description
Along Cedar River at road 40.3, north of the river. Field notes include an extensive plant list. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
22. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, King County:
Cascade Mountains of Western Washington: Eagle Ridge Creek Marsh.
Elev. 1620 ft. 47° 21' 42.1" N, 121° 37' 59.4" W; T22N R9E S27 NE 1/4.
Collected between road 100-300 and midway into the marsh, near east end of marsh. Growing on rotten log (decay class 3) in woods on south side of marsh. Phenology: Fertile. Origin: Native.
WTU 351425; Barcode: WTU-V-015501
Judy Harpel 01-2    26 May 2001
with Wayne Harpel, Rena Hamburger
22. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
24435 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
351425 
Barcode
WTU-V-015501 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Jennifer French, 23 2002.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Judy Harpel 
Collection #
01-2 
Other Collectors
Wayne Harpel, Rena Hamburger 
Date Collected
26 May 2001 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
King 
Locality
Cascade Mountains of Western Washington: Eagle Ridge Creek Marsh. 
Elevation
1620 ft. 
Coordinates
47° 21' 42.1" N, 121° 37' 59.4" W; T22N R9E S27 NE 1/4 
Site Description
Collected between road 100-300 and midway into the marsh, near east end of marsh. 
Specimen Notes
Growing on rotten log (decay class 3) in woods on south side of marsh. 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
23. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Whatcom County:
North Cascades National Park; Thunder Creek, 200-250 meters north of the suspension bridge across lower Thunder Creek.
Elev. 1350 ft. 48° 40' 42.16344" N, 121° 5' 14.25216" W; UTM Zone 10, 640800E, 5393250N. Coordinate Source: Calc. from UTM. UTM from field notes.
Old growth coniferous forest dominated by Pseudotsuga menziesii and Tsuga heterophylla; partial sun. On large decaying logs; partial sun. Phenology: Fertile. Origin: Native.
WTU 354120; Barcode: WTU-V-015415
NPS accession 632, catalog 23030
Steve Hahn 533    14 Jun 2002
with Mignonne Bivin
23. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
26554 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
354120 
Barcode
WTU-V-015415 
Other Numbers
NPS accession 632, catalog 23030 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ben Legler, 6 2002.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
Steve Hahn 
Collection #
533 
Other Collectors
Mignonne Bivin 
Date Collected
14 Jun 2002 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Whatcom 
Locality
North Cascades National Park; Thunder Creek, 200-250 meters north of the suspension bridge across lower Thunder Creek. 
Elevation
1350 ft. 
Coordinates
48° 40' 42.16344" N, 121° 5' 14.25216" W; UTM Zone 10, 640800E, 5393250N 
Site Description
Old growth coniferous forest dominated by Pseudotsuga menziesii and Tsuga heterophylla; partial sun. 
Specimen Notes
On large decaying logs; partial sun. 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
24. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Whatcom County:
North Cascades National Park; Perry Creek Shelter, near confluence of Perry Creek and Little Beaver Creek.
Elev. 2350 ft. 48° 55.2' N, 121° 8.9' W; UTM Zone 10, 635653.508E, 5419996.201N. Coordinate Source: Field notes. UTM calculated from Lat-Lon.
Shady forest of Thuja plicata and Pseudotsuga menziesii; thick duff, loose soil. Phenology: Fertile. Origin: Native.
WTU 353970; Barcode: WTU-V-015495
NPS accession 632, catalog 23951
Justin Hohn 17    13 Jul 2002
24. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
26637 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
353970 
Barcode
WTU-V-015495 
Other Numbers
NPS accession 632, catalog 23951 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by David Giblin, 26 2002.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Justin Hohn 
Collection #
17 
Other Collectors
 
Date Collected
13 Jul 2002 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Whatcom 
Locality
North Cascades National Park; Perry Creek Shelter, near confluence of Perry Creek and Little Beaver Creek. 
Elevation
2350 ft. 
Coordinates
48° 55.2' N, 121° 8.9' W; UTM Zone 10, 635653.508E, 5419996.201N 
Site Description
Shady forest of Thuja plicata and Pseudotsuga menziesii; thick duff, loose soil. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
25. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Whatcom County:
Little Beaver Trail, about 1 mile east of Perry Creek; North Cascades National Park.
Elev. 2034 ft. 48° 55.3' N, 121° 8.1' W; UTM Zone 10, 636625.715E, 5420205.349N. Coordinate Source: Field notes. UTM calculated from Lat-Lon.
Dry gneiss talus slope and adjacent forest of Pseudotsuga, with small cold creek; south aspect. Growing in part shade by creek. Phenology: Fertile. Origin: Native.
WTU 354463; Barcode: WTU-V-015629
NPS accession 632, catalog 24007
Ben Legler 28    13 Jul 2002
25. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
26981 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
354463 
Barcode
WTU-V-015629 
Other Numbers
NPS accession 632, catalog 24007 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by David Giblin, 26 2002.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
Ben Legler 
Collection #
28 
Other Collectors
 
Date Collected
13 Jul 2002 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Whatcom 
Locality
Little Beaver Trail, about 1 mile east of Perry Creek; North Cascades National Park. 
Elevation
2034 ft. 
Coordinates
48° 55.3' N, 121° 8.1' W; UTM Zone 10, 636625.715E, 5420205.349N 
Site Description
Dry gneiss talus slope and adjacent forest of Pseudotsuga, with small cold creek; south aspect. 
Specimen Notes
Growing in part shade by creek. 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
26. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Russian Federation, Sakhalin Region
Sakhalin Island; on southern part of island along the Nayba River circa 5 kilometers north-northwest of town of Bykov.
Elev. 98 ft. 47° 59' 54.48" N, 142° 53' 56.52" E.
Ecotone between uncut field along river and dense Abies Sachalinensis forest on slope above. Fronds clustered; in shade of Abies. Phenology: Fertile. Origin: Native.
WTU 357048
Ben Legler 680    17 Jul 2003
26. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
28056 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
357048 
Barcode
 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Slava Barkalov, 2003.  
Type Notes
 
Collector
Ben Legler 
Collection #
680 
Other Collectors
 
Date Collected
17 Jul 2003 
Verbatim Date
 
Country
Russian Federation 
State or Province
Sakhalin Region 
County
 
Locality
Sakhalin Island; on southern part of island along the Nayba River circa 5 kilometers north-northwest of town of Bykov. 
Elevation
98 ft. 
Coordinates
47° 59' 54.48" N, 142° 53' 56.52" E 
Site Description
Ecotone between uncut field along river and dense Abies Sachalinensis forest on slope above. 
Specimen Notes
Fronds clustered; in shade of Abies. 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
27. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Russian Federation, Sakhalin Region
Sakhalin Island, central part; 45 km ESE of Tymovskoye and 9 km SSW oc Chamginsky Pass, at mouth of a tributary to the Skalistaya River.
Elev. 1443 ft.
Calcareous roadcut above stream; forest above of Abies and Betula with dense moss understory. Fronds clustered in moss under Betula. Phenology: Fertile. Origin: Native.
WTU 357297
Ben Legler 997    28 Jul 2003
27. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
28081 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
357297 
Barcode
 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Slava Barkalov, 2003.  
Type Notes
 
Collector
Ben Legler 
Collection #
997 
Other Collectors
 
Date Collected
28 Jul 2003 
Verbatim Date
 
Country
Russian Federation 
State or Province
Sakhalin Region 
County
 
Locality
Sakhalin Island, central part; 45 km ESE of Tymovskoye and 9 km SSW oc Chamginsky Pass, at mouth of a tributary to the Skalistaya River. 
Elevation
1443 ft. 
Coordinates
 
Site Description
Calcareous roadcut above stream; forest above of Abies and Betula with dense moss understory. 
Specimen Notes
Fronds clustered in moss under Betula. 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
28. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Pacific County:
The Nature Conservancy's Ellsworth Creek Preserve. Waypoint #36. Along trail in forest tract just above logging spur road.
Elev. 161 ft. 46° 23.68' N, 123° 53.054' W.
West of spur road on trail leading down to Ellsworth Creek. Phenology: Fertile. Origin: Native.
WTU 357525; Barcode: WTU-V-015481
Richard G. Olmstead 2003-46    31 May 2003
28. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
28178 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
357525 
Barcode
WTU-V-015481 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Wendy Gibble, 20 2004.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ben Legler, 18 2005. Sources: FNA Vol. 2.  
Type Notes
 
Collector
Richard G. Olmstead 
Collection #
2003-46 
Other Collectors
 
Date Collected
31 May 2003 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Pacific 
Locality
The Nature Conservancy's Ellsworth Creek Preserve. Waypoint #36. Along trail in forest tract just above logging spur road. 
Elevation
161 ft. 
Coordinates
46° 23.68' N, 123° 53.054' W 
Site Description
West of spur road on trail leading down to Ellsworth Creek. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
29. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Valdez Cordova County:
Evans Island. Port San Juan.
60.051325°, -148.071491°. Coordinate uncertainty: 1200 meters. Georeferenced by Joseph Stormer.
Shady stream bank. Origin: Native.
WTU 291662; Barcode: WTU-V-015461
W. J. Eyerdam 7138    25 Aug 1948
29. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31739 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
291662 
Barcode
WTU-V-015461 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by W.J.E..
2. Dryopteris austriaca (Jacq.) Woynar var. austriaca.
    Annotated by C. V. Morton, 1954. Remarks: U.S. National Museum.
3. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by F.W. Crane, 1957.
4. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
W. J. Eyerdam 
Collection #
7138 
Other Collectors
 
Date Collected
25 Aug 1948 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Valdez Cordova 
Locality
Evans Island. Port San Juan. 
Elevation
 
Coordinates
60.051325°, -148.071491° 
Site Description
Shady stream bank. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
30. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Kodiak Island County:
Port Vita, Raspberry Strait, Raspberry Island, Kodiak group.
Elev. 1968 ft. 58.065556°, -153.076792°. Coordinate uncertainty: 3000 meters. Georeferenced by Joseph Stormer.
Spruce forest. Origin: Native.
WTU 291663; Barcode: WTU-V-015463
W. J. Eyerdam 4028     1945
30. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31750 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
291663 
Barcode
WTU-V-015463 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris spinulosa (O.F. Muell.) Kunze.
    Annotated by W. J. Eyerdam, 3 1945.
2. Dryopteris austriaca (Jacq.) Woynar var. austriaca.
    Annotated by C. V. Morton, 1954. Remarks: U.S. National Museum.
3. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by F. W. Crane, 1957.
4. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
W. J. Eyerdam 
Collection #
4028 
Other Collectors
 
Date Collected
1945 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Kodiak Island 
Locality
Port Vita, Raspberry Strait, Raspberry Island, Kodiak group. 
Elevation
1968 ft. 
Coordinates
58.065556°, -153.076792° 
Site Description
Spruce forest. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
31. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Juneau County:
(locality not given)
Woods. Origin: Native.
WTU 293125; Barcode: WTU-V-015462
J. P. Anderson 6144    28 Jul 1940
31. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31755 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
293125 
Barcode
WTU-V-015462 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar var. austriaca.
    Annotated by C. V. Morton, 1954. Remarks: U.S. National Museum.
3. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by F. W. Crane, 1957.
4. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
J. P. Anderson 
Collection #
6144 
Other Collectors
 
Date Collected
28 Jul 1940 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Juneau 
Locality
(locality not given) 
Elevation
 
Coordinates
 
Site Description
Woods. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
32. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Wrangell Petersburg County:
Washington Bay, Kuiu Island.
56.71885°, -134.364815°. Datum: WGS 84. Coordinate uncertainty: 2000 meters. Georeferenced by Joseph Stormer. Coordinate Source: Digital Map.
Shady spruce and hemlock forest. Change in collection number 8325=>8328. Origin: Native.
WTU 294342; Barcode: WTU-V-015464
W. J. Eyerdam 8328    19 Aug 1956
32. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31759 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
294342 
Barcode
WTU-V-015464 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by W.J.E..
2. Dryopteris austriaca (Jacq.) Woynar.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
W. J. Eyerdam 
Collection #
8328 
Other Collectors
 
Date Collected
19 Aug 1956 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Wrangell Petersburg 
Locality
Washington Bay, Kuiu Island. 
Elevation
 
Coordinates
56.71885°, -134.364815° 
Site Description
Shady spruce and hemlock forest. 
Specimen Notes
Change in collection number 8325=>8328. 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
33. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Wrangell Petersburg County:
Washington Bay, Kuiu Island.
56.71885°, -134.364815°. Datum: WGS 84. Coordinate uncertainty: 2000 meters. Georeferenced by Joseph Stormer. Coordinate Source: Digital Map.
Spruce and hemlock forest. Origin: Native.
WTU 294341; Barcode: WTU-V-015465
W. J. Eyerdam 8249    25 Jul 1956
33. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31761 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
294341 
Barcode
WTU-V-015465 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by W. J. Eyerdam. Remarks: crossed out.
2. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by W.J.E..
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
W. J. Eyerdam 
Collection #
8249 
Other Collectors
 
Date Collected
25 Jul 1956 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Wrangell Petersburg 
Locality
Washington Bay, Kuiu Island. 
Elevation
 
Coordinates
56.71885°, -134.364815° 
Site Description
Spruce and hemlock forest. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
34. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Sitka County:
Southeast, Tongass National Forest, Baranof Island, Takaz Bay [Takatz Bay], just east of tide flats on small treeless dome shape islet.
57.149167°, -134.826944°. Datum: WGS 84. Coordinate uncertainty: 301 meters. Georeferenced by Joseph Stormer. Coordinate Source: Digital Map.
Islet, few pockets of soil, largest plant Sambucus Racemosa. Origin: Native.
WTU 282584; Barcode: WTU-V-015460
Mary Clay Muller 3903    29 Jul 1980
34. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31764 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
282584 
Barcode
WTU-V-015460 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by M. Muller, 1980.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
Mary Clay Muller 
Collection #
3903 
Other Collectors
 
Date Collected
29 Jul 1980 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Sitka 
Locality
Southeast, Tongass National Forest, Baranof Island, Takaz Bay [Takatz Bay], just east of tide flats on small treeless dome shape islet. 
Elevation
 
Coordinates
57.149167°, -134.826944° 
Site Description
Islet, few pockets of soil, largest plant Sambucus Racemosa. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
35. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Juneau County:
Southeast, mainland, east of Juneau, Mount Roberts Trail, lower part of the trail in the woods.
Elev. 500 ft.
Woods. More robust than usual. Origin: Native.
WTU 289056; Barcode: WTU-V-015459
Mary Clay Muller 3391    11 Aug 1979
35. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31767 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
289056 
Barcode
WTU-V-015459 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
Mary Clay Muller 
Collection #
3391 
Other Collectors
 
Date Collected
11 Aug 1979 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Juneau 
Locality
Southeast, mainland, east of Juneau, Mount Roberts Trail, lower part of the trail in the woods. 
Elevation
500 ft. 
Coordinates
 
Site Description
Woods. 
Specimen Notes
More robust than usual. 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
36. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Juneau County:
Southeast, mainland, east of Juneau, Mount Roberts Trail, lower part of the trail in the woods.
Elev. 500 ft.
Woods. Origin: Native.
WTU 289055; Barcode: WTU-V-015458
Mary Clay Muller 3389    11 Aug 1979
36. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31769 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
289055 
Barcode
WTU-V-015458 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by M. Muller, 1979.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
Mary Clay Muller 
Collection #
3389 
Other Collectors
 
Date Collected
11 Aug 1979 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Juneau 
Locality
Southeast, mainland, east of Juneau, Mount Roberts Trail, lower part of the trail in the woods. 
Elevation
500 ft. 
Coordinates
 
Site Description
Woods. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
37. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Kodiak Island County:
Olga Bay, lower mountain slope, in tundra.
Tundra. Spore collected 7 out 80. Origin: Native.
WTU 45855; Barcode: WTU-V-015457
Ethel H. Looff X40    2 Aug 1938
with Henry B. Looff
37. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31771 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
45855 
Barcode
WTU-V-015457 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by Dr. William R. Maxon.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
Ethel H. Looff 
Collection #
X40 
Other Collectors
Henry B. Looff 
Date Collected
2 Aug 1938 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Kodiak Island 
Locality
Olga Bay, lower mountain slope, in tundra. 
Elevation
 
Coordinates
 
Site Description
Tundra. 
Specimen Notes
Spore collected 7 out 80. 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
38. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Kodiak Island County:
Olga Bay.
Valley. 7 out of 80 spores collected. Origin: Native.
WTU 45853; Barcode: WTU-V-015456
Ethel H. Looff X52     Aug 1938
with Henry B. Looff
38. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31773 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
45853 
Barcode
WTU-V-015456 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by Dr. William R. Maxon.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
Ethel H. Looff 
Collection #
X52 
Other Collectors
Henry B. Looff 
Date Collected
Aug 1938 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Kodiak Island 
Locality
Olga Bay. 
Elevation
 
Coordinates
 
Site Description
Valley. 
Specimen Notes
7 out of 80 spores collected. 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
39. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Nome County:
Seward Peninsula, Cape Nome.
64.473659°, -164.997849°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by Joseph Stormer. Coordinate Source: Digital Map.
Rocky hillside. Origin: Native.
WTU 12298JWT; Barcode: WTU-V-015454
George Neville Jones 8961     Jul 1936
39. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31780 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
12298JWT 
Barcode
WTU-V-015454 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris hastata (Fée) Urb.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946. Remarks: Herbarium of the University of California.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
George Neville Jones 
Collection #
8961 
Other Collectors
 
Date Collected
Jul 1936 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Nome 
Locality
Seward Peninsula, Cape Nome. 
Elevation
 
Coordinates
64.473659°, -164.997849° 
Site Description
Rocky hillside. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
40. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Nome County:
Seward Peninsula, Cape Nome.
64.473659°, -164.997849°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by Joseph Stormer. Coordinate Source: Digital Map.
Rocky hillside. Origin: Native.
WTU 37654; Barcode: WTU-V-015453
George Neville Jones 8961     Jul 1936
40. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31781 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
37654 
Barcode
WTU-V-015453 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris aquilonaris.
    Remarks: Crossed out.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1945. Remarks: D. aquilonaris is a form of D. fragraus.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
George Neville Jones 
Collection #
8961 
Other Collectors
 
Date Collected
Jul 1936 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Nome 
Locality
Seward Peninsula, Cape Nome. 
Elevation
 
Coordinates
64.473659°, -164.997849° 
Site Description
Rocky hillside. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
41. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Nome County:
Seward Peninsula, Cape Nome.
64.473659°, -164.997849°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by Joseph Stormer. Coordinate Source: Digital Map.
Rocky hillside. Origin: Native.
WTU 12275JWT; Barcode: WTU-V-015452
George Neville Jones 8961     Jul 1936
41. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31783 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
12275JWT 
Barcode
WTU-V-015452 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris hastata (Fée) Urb.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
George Neville Jones 
Collection #
8961 
Other Collectors
 
Date Collected
Jul 1936 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Nome 
Locality
Seward Peninsula, Cape Nome. 
Elevation
 
Coordinates
64.473659°, -164.997849° 
Site Description
Rocky hillside. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
42. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Juneau County:
Juneau.
58.301944°, -134.419722°. Coordinate uncertainty: 10000 meters.
Origin: Native.
WTU 90301; Barcode: WTU-V-015451
Emma A. Shumway s.n.    23 Jul 1891
42. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31786 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
90301 
Barcode
WTU-V-015451 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris hastata (Fée) Urb.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
Emma A. Shumway 
Collection #
s.n. 
Other Collectors
 
Date Collected
23 Jul 1891 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Juneau 
Locality
Juneau. 
Elevation
 
Coordinates
58.301944°, -134.419722° 
Site Description
 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
43. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Kodiak Island County:
Olga Bay, Lower mountain slopes.
Origin: Native.
WTU 61401; Barcode: WTU-V-015450
Ethel H. Looff X58     Aug 1938
with Henry B. Looff
43. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31787 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
61401 
Barcode
WTU-V-015450 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
Ethel H. Looff 
Collection #
X58 
Other Collectors
Henry B. Looff 
Date Collected
Aug 1938 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Kodiak Island 
Locality
Olga Bay, Lower mountain slopes. 
Elevation
 
Coordinates
 
Site Description
 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
44. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Aleutians West County:
Chicagof Harbor, at foot of hill east of Attu Village site.
52.924031°, 173.253523°. Datum: WGS 84. Coordinate uncertainty: 1219 meters. Georeferenced by Joseph Stormer. Coordinate Source: Digital Map.
Origin: Native.
WTU 155643; Barcode: WTU-V-015449
Lt. G. B. Schaack 679    24 Jul 1945
44. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31789 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
155643 
Barcode
WTU-V-015449 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by W. R. Maxon.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
Lt. G. B. Schaack 
Collection #
679 
Other Collectors
 
Date Collected
24 Jul 1945 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Aleutians West 
Locality
Chicagof Harbor, at foot of hill east of Attu Village site. 
Elevation
 
Coordinates
52.924031°, 173.253523° 
Site Description
 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
45. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Kodiak Island County:
Olga Bay, Lower mountain slopes.
Origin: Native.
WTU 23356JWT; Barcode: WTU-V-015448
Ethel H. Looff X57     Aug 1938
with Henry B. Looff
45. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31792 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
23356JWT 
Barcode
WTU-V-015448 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
Ethel H. Looff 
Collection #
X57 
Other Collectors
Henry B. Looff 
Date Collected
Aug 1938 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Kodiak Island 
Locality
Olga Bay, Lower mountain slopes. 
Elevation
 
Coordinates
 
Site Description
 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
46. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Wrangell Petersburg County:
Washington Bay, Kuiu Island.
56.71885°, -134.364815°. Datum: WGS 84. Coordinate uncertainty: 2000 meters. Georeferenced by Joseph Stormer. Coordinate Source: Digital Map.
Deep shady spruce forest. Origin: Native.
WTU 118036; Barcode: WTU-V-015447
W. J. Eyerdam 5450    20 Aug 1947
46. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31793 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
118036 
Barcode
WTU-V-015447 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by W.J.E..
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
W. J. Eyerdam 
Collection #
5450 
Other Collectors
 
Date Collected
20 Aug 1947 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Wrangell Petersburg 
Locality
Washington Bay, Kuiu Island. 
Elevation
 
Coordinates
56.71885°, -134.364815° 
Site Description
Deep shady spruce forest. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
47. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Kodiak Island County:
Port Vita, Raspberry Strait, Raspberry Island, Kodiak group.
Elev. 1968 ft. 58.065556°, -153.076792°. Coordinate uncertainty: 3000 meters. Georeferenced by Joseph Stormer.
Origin: Native.
WTU 190893; Barcode: WTU-V-015446
W. J. Eyerdam 4028     1945
47. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31795 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
190893 
Barcode
WTU-V-015446 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris spinulosa (O.F. Muell.) Kunze.
    Annotated by W.J.E., 3 1945.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
W. J. Eyerdam 
Collection #
4028 
Other Collectors
 
Date Collected
1945 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Kodiak Island 
Locality
Port Vita, Raspberry Strait, Raspberry Island, Kodiak group. 
Elevation
1968 ft. 
Coordinates
58.065556°, -153.076792° 
Site Description
 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
48. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Nome County:
Base of mountains and slopes, 16 miles west of Nome City.
Grassy. Origin: Native.
WTU 36980; Barcode: WTU-V-015445
J. B. Flett 1506    20 Jul 1900
48. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31797 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
36980 
Barcode
WTU-V-015445 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris aquilonaris.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 12 1945. Remarks: D. aquilonaris is a form of D. fragraus.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
J. B. Flett 
Collection #
1506 
Other Collectors
 
Date Collected
20 Jul 1900 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Nome 
Locality
Base of mountains and slopes, 16 miles west of Nome City. 
Elevation
 
Coordinates
 
Site Description
Grassy. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
49. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Valdez Cordova County:
Middleton Island is located in the Gulf of Alaska about 115 miles south of Valdez.
Steep west facing bluff. Common. Origin: Native.
WTU 181718; Barcode: WTU-V-015437
John H. Thomas 5813    6 Jun 1956
49. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31798 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
181718 
Barcode
WTU-V-015437 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
John H. Thomas 
Collection #
5813 
Other Collectors
 
Date Collected
6 Jun 1956 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Valdez Cordova 
Locality
Middleton Island is located in the Gulf of Alaska about 115 miles south of Valdez. 
Elevation
 
Coordinates
 
Site Description
Steep west facing bluff. 
Specimen Notes
Common. 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
50. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Valdez Cordova County:
Thum Bay, Knight Island, Prince William Sound.
60.206643°, -147.811189°. Coordinate uncertainty: 2400 meters.
Deep moss in spruce forest. Origin: Native.
WTU 291665; Barcode: WTU-V-015444
W. J. Eyerdam 3634    20 Sep 1940
50. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31800 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
291665 
Barcode
WTU-V-015444 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris spinulosa (O.F. Muell.) Kunze.
    Annotated by W.J.E..
2. Dryopteris austriaca (Jacq.) Woynar.
3. Dryopteris austriaca (Jacq.) Woynar var. austriaca.
    Annotated by C. V. Morton, 1954. Remarks: U.S. National Museum.
4. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by F. W. Crane, 1957.
5. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
W. J. Eyerdam 
Collection #
3634 
Other Collectors
 
Date Collected
20 Sep 1940 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Valdez Cordova 
Locality
Thum Bay, Knight Island, Prince William Sound. 
Elevation
 
Coordinates
60.206643°, -147.811189° 
Site Description
Deep moss in spruce forest. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
51. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Wrangell Petersburg County:
Washington Bay, Kuiu Island.
56.71885°, -134.364815°. Datum: WGS 84. Coordinate uncertainty: 2000 meters. Georeferenced by Joseph Stormer. Coordinate Source: Digital Map.
Deep shady spruce forest. Origin: Native.
WTU 291664; Barcode: WTU-V-015441
W. J. Eyerdam 5450    20 Aug 1947
51. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31802 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
291664 
Barcode
WTU-V-015441 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by W.J.E..
2. Dryopteris austriaca (Jacq.) Woynar var. austriaca.
    Annotated by C. V. Morton, 1954. Remarks: U.S. National Museum.
3. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by F. W. Crane, 1957.
4. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006.  
Type Notes
 
Collector
W. J. Eyerdam 
Collection #
5450 
Other Collectors
 
Date Collected
20 Aug 1947 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Wrangell Petersburg 
Locality
Washington Bay, Kuiu Island. 
Elevation
 
Coordinates
56.71885°, -134.364815° 
Site Description
Deep shady spruce forest. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
52. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Snohomish County:
Marysville.
48.05194°, -122.17583°. Datum: WGS 84. Coordinate uncertainty: 3570 meters. Georeferenced by Maria Yousoufian. Coordinate Source: GeoLocate.
Shady woods. Phenology: Fertile. Origin: Native.
WTU 43907; Barcode: WTU-V-015594
J. M. Grant s.n.     Jun 1929
52. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31803 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
43907 
Barcode
WTU-V-015594 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris spinulosa (O.F. Muell.) Watt var. dilatata auct. non (Hoffmann) Underwood.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
J. M. Grant 
Collection #
s.n. 
Other Collectors
 
Date Collected
Jun 1929 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Snohomish 
Locality
Marysville. 
Elevation
 
Coordinates
48.05194°, -122.17583° 
Site Description
Shady woods. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
53. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Valdez Cordova County:
Thum Bay, Knight Island, Prince William Sound.
Elev. 328 ft. 60.206643°, -147.811189°. Coordinate uncertainty: 2400 meters.
Shade of overhanging cliff amongst moss. Origin: Native.
WTU 291669; Barcode: WTU-V-015440
Walter J. Eyerdam 3300    2 Oct 1939
53. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31804 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
291669 
Barcode
WTU-V-015440 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris spinulosa (O.F. Muell.) Kunze.
    Annotated by L. Constance.
2. Dryopteris austriaca (Jacq.) Woynar var. austriaca.
    Annotated by C. V. Morton, 1954. Remarks: U.S. National Museum.
3. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by F. W. Crane, 1957.
4. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
Walter J. Eyerdam 
Collection #
3300 
Other Collectors
 
Date Collected
2 Oct 1939 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Valdez Cordova 
Locality
Thum Bay, Knight Island, Prince William Sound. 
Elevation
328 ft. 
Coordinates
60.206643°, -147.811189° 
Site Description
Shade of overhanging cliff amongst moss. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
54. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Snohomish County:
Mount Pugh, Cascade Mountains.
Elev. 7000 ft. 48.143056°, -121.372778°. Datum: NAD 27. Coordinate uncertainty: 1000 meters. Georeferenced by WTU Staff. Coordinate Source: Georeferenced.
Wooded slopes. Phenology: Fertile. Origin: Native.
WTU 17602JWT; Barcode: WTU-V-015595
J. William Thompson 14356    18 Aug 1938
54. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31806 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
17602JWT 
Barcode
WTU-V-015595 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 14 2004. Remarks: WTU. 
Type Notes
 
Collector
J. William Thompson 
Collection #
14356 
Other Collectors
 
Date Collected
18 Aug 1938 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Snohomish 
Locality
Mount Pugh, Cascade Mountains. 
Elevation
7000 ft. 
Coordinates
48.143056°, -121.372778° 
Site Description
Wooded slopes. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
55. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Jefferson County:
Dosewallips River.
47.688764°, -122.897951°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Damp rich woods. Phenology: Fertile. Origin: Native.
WTU 39478; Barcode: WTU-V-015596
G. N. Jones 5851    8 Aug 1934
55. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31807 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
39478 
Barcode
WTU-V-015596 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
G. N. Jones 
Collection #
5851 
Other Collectors
 
Date Collected
8 Aug 1934 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Jefferson 
Locality
Dosewallips River. 
Elevation
 
Coordinates
47.688764°, -122.897951° 
Site Description
Damp rich woods. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
56. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Wrangell Petersburg County:
Washington Bay, Kuiu Island.
56.71885°, -134.364815°. Datum: WGS 84. Coordinate uncertainty: 2000 meters. Georeferenced by Joseph Stormer. Coordinate Source: Digital Map.
Shady spruce and hemlock forest. Collection # changed 8325=>8328. Origin: Native.
WTU 294345; Barcode: WTU-V-015439
W. J. Eyerdam 8328    19 Aug 1956
56. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31808 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
294345 
Barcode
WTU-V-015439 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by W.J.E.. Remarks: #NAME?
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.  
Type Notes
 
Collector
W. J. Eyerdam 
Collection #
8328 
Other Collectors
 
Date Collected
19 Aug 1956 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Wrangell Petersburg 
Locality
Washington Bay, Kuiu Island. 
Elevation
 
Coordinates
56.71885°, -134.364815° 
Site Description
Shady spruce and hemlock forest. 
Specimen Notes
Collection # changed 8325=>8328. 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
57. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Clallam County:
Lower slope of Mt. Angeles.
Elev. 3000 ft. 47.995278°, -123.4575°. Datum: NAD 27. Coordinate uncertainty: 5000 meters. Georeferenced by WTU Staff. Coordinate Source: Georeferenced.
Wooded slope. Phenology: Fertile. Origin: Native.
WTU 3197JWT; Barcode: WTU-V-015587
J. William Thompson 5506    2 Aug 1930
57. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31810 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
3197JWT 
Barcode
WTU-V-015587 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
J. William Thompson 
Collection #
5506 
Other Collectors
 
Date Collected
2 Aug 1930 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Clallam 
Locality
Lower slope of Mt. Angeles. 
Elevation
3000 ft. 
Coordinates
47.995278°, -123.4575° 
Site Description
Wooded slope. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
58. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Aleutians West County:
Aleutians Island, Engineer Canyon, Siddens Valley.
Elev. 250 ft. 52.884051°, 173.193541°. Datum: WGS 84. Coordinate uncertainty: 400 meters. Georeferenced by Joseph Stormer. Coordinate Source: Digital Map.
Steep slope among tall herbs and ferns. Origin: Native.
WTU 291583; Barcode: WTU-V-015436
Robert M. Hardy 318    8 Aug 1945
58. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31811 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
291583 
Barcode
WTU-V-015436 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
    Remarks: Crossed out.
2. Dryopteris dilatata auct. non (Hoffm.n) Gray.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006.  
Type Notes
 
Collector
Robert M. Hardy 
Collection #
318 
Other Collectors
 
Date Collected
8 Aug 1945 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Aleutians West 
Locality
Aleutians Island, Engineer Canyon, Siddens Valley. 
Elevation
250 ft. 
Coordinates
52.884051°, 173.193541° 
Site Description
Steep slope among tall herbs and ferns. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
59. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Clallam County:
Lower slopes of Mt. Angeles, Olympic Mountains.
Elev. 2000 ft. 47.995278°, -123.4575°. Georeferenced by Ed Alverson. Coordinate Source: Inferred from duplicate specimen.
Wooded slopes. Phenology: Fertile. Origin: Native.
WTU 3226JWT; Barcode: WTU-V-015597
J. William Thompson 7561    20 Jul 1931
59. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31812 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
3226JWT 
Barcode
WTU-V-015597 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Thelypteris spinulosa (O.F.Muell.) Nieuwl. var. dilatata (Hoffm.)St.John.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
J. William Thompson 
Collection #
7561 
Other Collectors
 
Date Collected
20 Jul 1931 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Clallam 
Locality
Lower slopes of Mt. Angeles, Olympic Mountains. 
Elevation
2000 ft. 
Coordinates
47.995278°, -123.4575° 
Site Description
Wooded slopes. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
60. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Alaska, Kodiak Island County:
Port Vita, Raspberry Island.
Elev. 32 ft. 58.065556°, -153.076792°. Coordinate uncertainty: 3000 meters. Georeferenced by Joseph Stormer.
In rich woods under alnus bushes. Origin: Native.
WTU 291668; Barcode: WTU-V-015443
W. J. Eyerdam 3339    20 Aug 1939
60. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31813 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
291668 
Barcode
WTU-V-015443 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
2. Dryopteris austriaca (Jacq.) Woynar var. austriaca.
    Annotated by C. V. Morton, 1954. Remarks: U.S. National Museum.
3. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by F. W. Crane, 1957.
4. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by E. Alverson, 1 2006. Sources: FNA.
4. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA.
5. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 31 2006. Sources: FNA.  
Type Notes
 
Collector
W. J. Eyerdam 
Collection #
3339 
Other Collectors
 
Date Collected
20 Aug 1939 
Verbatim Date
 
Country
U.S.A. 
State or Province
Alaska 
County
Kodiak Island 
Locality
Port Vita, Raspberry Island. 
Elevation
32 ft. 
Coordinates
58.065556°, -153.076792° 
Site Description
In rich woods under alnus bushes. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
61. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, King County:
At Snoqualmie Pass. Cascade Mountains.
Elev. 3010 ft. 47.52889°, -121.82417°. Datum: WGS 84. Coordinate uncertainty: 5000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Base of shaded talus. Phenology: Fertile. Origin: Native.
WTU 3348JWT; Barcode: WTU-V-015598
J. William Thompson 8924    17 Sep 1932
61. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31814 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
3348JWT 
Barcode
WTU-V-015598 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Thelypteris dilatata (Hoffm.)House.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
J. William Thompson 
Collection #
8924 
Other Collectors
 
Date Collected
17 Sep 1932 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
King 
Locality
At Snoqualmie Pass. Cascade Mountains. 
Elevation
3010 ft. 
Coordinates
47.52889°, -121.82417° 
Site Description
Base of shaded talus. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
62. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Clackamas County:
Boring.
Origin: Native.
WTU 272659; Barcode: WTU-V-015420
Wilhelm N. Suksdorf 2030    20 Aug 1903
62. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31815 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
272659 
Barcode
WTU-V-015420 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
Wilhelm N. Suksdorf 
Collection #
2030 
Other Collectors
 
Date Collected
20 Aug 1903 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Clackamas 
Locality
Boring. 
Elevation
 
Coordinates
 
Site Description
 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
63. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Pierce County:
Trail to Van Trump Park, Mt. Rainier.
Elev. 5000 ft. 46.79806°, -121.77806°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Alpine rock slide under alders. Phenology: Fertile. Origin: Native.
WTU 33037; Barcode: WTU-V-015599
J. William Thompson 12556    14 Aug 1935
63. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31817 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
33037 
Barcode
WTU-V-015599 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
J. William Thompson 
Collection #
12556 
Other Collectors
 
Date Collected
14 Aug 1935 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Pierce 
Locality
Trail to Van Trump Park, Mt. Rainier. 
Elevation
5000 ft. 
Coordinates
46.79806°, -121.77806° 
Site Description
Alpine rock slide under alders. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
64. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Jefferson County:
Along Hoh River near Jackson Ranger Station.
47.814011°, -124.176698°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Phenology: Spores. Origin: Native.
WTU 28285; Barcode: WTU-V-015600
I. C. Otis 1364    6 Aug 1924
64. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31818 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
28285 
Barcode
WTU-V-015600 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946. Remarks: [D. dilatata (Hoffmann) A. Gray] Herbarium of the University of California.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
I. C. Otis 
Collection #
1364 
Other Collectors
 
Date Collected
6 Aug 1924 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Jefferson 
Locality
Along Hoh River near Jackson Ranger Station. 
Elevation
 
Coordinates
47.814011°, -124.176698° 
Site Description
 
Specimen Notes
 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
65. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Clallam County:
Lake Crescent.
48.09472°, -123.80583°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Phenology: Fertile. Origin: Native.
WTU 21392; Barcode: WTU-V-015601
Geo. N. Jones 3431     Jul 1931
65. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31820 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
21392 
Barcode
WTU-V-015601 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Thelypteris spinulosa (O.F.Muell.) Nieuwl. var. dilatata (Hoffm.)St.John.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Geo. N. Jones 
Collection #
3431 
Other Collectors
 
Date Collected
Jul 1931 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Clallam 
Locality
Lake Crescent. 
Elevation
 
Coordinates
48.09472°, -123.80583° 
Site Description
 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
66. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Curry County:
Gold Beach.
Origin: Native.
WTU 3351JWT; Barcode: WTU-V-015419
Ira Gabrielson s.n.     1927
66. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31821 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
3351JWT 
Barcode
WTU-V-015419 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
Ira Gabrielson 
Collection #
s.n. 
Other Collectors
 
Date Collected
1927 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Curry 
Locality
Gold Beach. 
Elevation
 
Coordinates
 
Site Description
 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
67. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Island County:
Cranberry Lake, Deception Pass State Park, Whidbey Island.
48.14389°, -122.49111°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Phenology: Fertile. Origin: Native.
WTU 38857; Barcode: WTU-V-015602
Harold W. Smith 805    3 Jun 1936
67. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31824 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
38857 
Barcode
WTU-V-015602 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Harold W. Smith 
Collection #
805 
Other Collectors
 
Date Collected
3 Jun 1936 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Island 
Locality
Cranberry Lake, Deception Pass State Park, Whidbey Island. 
Elevation
 
Coordinates
48.14389°, -122.49111° 
Site Description
 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
68. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Tillamook County:
Neskowin Creek.
Origin: Native.
WTU 28006; Barcode: WTU-V-015411
G. E. Morrill 224    25 Jul 1933
68. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31825 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
28006 
Barcode
WTU-V-015411 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946. Remarks: The Herbarium of The University of California.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
G. E. Morrill 
Collection #
224 
Other Collectors
 
Date Collected
25 Jul 1933 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Tillamook 
Locality
Neskowin Creek. 
Elevation
 
Coordinates
 
Site Description
 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
69. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, San Juan County:
Friday Harbor.
48.541389°, -123.0225°. Datum: WGS 84. Coordinate uncertainty: 2000 meters. Georeferenced by WTU Staff. Coordinate Source: Georeferenced.
Phenology: Fertile. Origin: Native.
WTU 124826; Barcode: WTU-V-015603
A. S. Pope s.n.    25 Jun 1904
69. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31827 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
124826 
Barcode
WTU-V-015603 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
A. S. Pope 
Collection #
s.n. 
Other Collectors
 
Date Collected
25 Jun 1904 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
San Juan 
Locality
Friday Harbor. 
Elevation
 
Coordinates
48.541389°, -123.0225° 
Site Description
 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
70. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Tillamook County:
Head waters of the Wilson River, Coast Mountains.
Elev. 3000 ft.
Woods. Origin: Native.
WTU 3337JWT; Barcode: WTU-V-015409
J. William Thompson 3022    19 Jul 1927
70. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31828 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
3337JWT 
Barcode
WTU-V-015409 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946. Remarks: Herbarium of The University of California.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
J. William Thompson 
Collection #
3022 
Other Collectors
 
Date Collected
19 Jul 1927 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Tillamook 
Locality
Head waters of the Wilson River, Coast Mountains. 
Elevation
3000 ft. 
Coordinates
 
Site Description
Woods. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
71. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, King County:
10 miles from Renton.
47.48306°, -122.21583°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Moist woods. Phenology: Spores. Origin: Native.
WTU 31228; Barcode: WTU-V-015604
G. L. Wittrock s.n.    6 Apr 1928
71. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31829 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
31228 
Barcode
WTU-V-015604 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004. Remarks: WTU. 
Type Notes
 
Collector
G. L. Wittrock 
Collection #
s.n. 
Other Collectors
 
Date Collected
6 Apr 1928 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
King 
Locality
10 miles from Renton. 
Elevation
 
Coordinates
47.48306°, -122.21583° 
Site Description
Moist woods. 
Specimen Notes
 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
72. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Washington County:
Near Roderick Falls, near Forest Grove.
Deep ravine. Origin: Native.
WTU 3270JWT; Barcode: WTU-V-015418
J. William Thompson 4279a    26 May 1928
72. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31832 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
3270JWT 
Barcode
WTU-V-015418 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946. Remarks: Herbarium of The University of California.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
J. William Thompson 
Collection #
4279a 
Other Collectors
 
Date Collected
26 May 1928 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Washington 
Locality
Near Roderick Falls, near Forest Grove. 
Elevation
 
Coordinates
 
Site Description
Deep ravine. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
73. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Clallam County:
Canyon Creek, Olympic National Forest.
Elev. 2100 ft. 48.00752°, -124.093917°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Tsuga heterophylla-Abines amabilis forest. Canadian zone. Phenology: Vegetative. Origin: Native.
WTU 30288; Barcode: WTU-V-015619
George Neville Jones 8364    24 Aug 1935
73. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31834 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
30288 
Barcode
WTU-V-015619 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006.  
Type Notes
 
Collector
George Neville Jones 
Collection #
8364 
Other Collectors
 
Date Collected
24 Aug 1935 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Clallam 
Locality
Canyon Creek, Olympic National Forest. 
Elevation
2100 ft. 
Coordinates
48.00752°, -124.093917° 
Site Description
Tsuga heterophylla-Abines amabilis forest. Canadian zone. 
Specimen Notes
 
Phenology
Vegetative 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
74. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Clatsop County:
Warrenton.
Origin: Native.
WTU 197116; Barcode: WTU-V-015410
Grace M. Cole 15    28 Aug 1932
74. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31835 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
197116 
Barcode
WTU-V-015410 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Thelypteris spinulosa (O.F.Muell.) Nieuwl. var. dilatata (Hoffm.)St.John.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
Grace M. Cole 
Collection #
15 
Other Collectors
 
Date Collected
28 Aug 1932 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Clatsop 
Locality
Warrenton. 
Elevation
 
Coordinates
 
Site Description
 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
75. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Snohomish County:
Mount Pugh, Cascade Mountains.
Elev. 7000 ft. 48.143056°, -121.372778°. Datum: NAD 27. Coordinate uncertainty: 1000 meters. Georeferenced by WTU Staff. Coordinate Source: Georeferenced.
Wooded slopes. Phenology: Fertile. Origin: Native.
WTU 49057; Barcode: WTU-V-015593
J. William Thompson 14356    18 Aug 1938
75. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31837 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
49057 
Barcode
WTU-V-015593 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
J. William Thompson 
Collection #
14356 
Other Collectors
 
Date Collected
18 Aug 1938 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Snohomish 
Locality
Mount Pugh, Cascade Mountains. 
Elevation
7000 ft. 
Coordinates
48.143056°, -121.372778° 
Site Description
Wooded slopes. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
76. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Baker County:
Above Cornucopia, Wallowa Mountains.
Elev. 7000 ft.
Alpine stream in brush. Origin: Native.
WTU 3022JWT; Barcode: WTU-V-015427
J. William Thompson 13413    20 Jul 1936
76. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31839 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
3022JWT 
Barcode
WTU-V-015427 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946. Remarks: Herbarium of The University of California.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
J. William Thompson 
Collection #
13413 
Other Collectors
 
Date Collected
20 Jul 1936 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Baker 
Locality
Above Cornucopia, Wallowa Mountains. 
Elevation
7000 ft. 
Coordinates
 
Site Description
Alpine stream in brush. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
77. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Mason County:
Twanoh State Park, Hood Canal.
47.37345°, -122.97428°. Datum: WGS 84. Coordinate uncertainty: 1000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Springy area. Phenology: Fertile. Origin: Native.
WTU 38664; Barcode: WTU-V-015588
Harold W. Smith 185    2 Feb 1936
77. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31841 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
38664 
Barcode
WTU-V-015588 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Harold W. Smith 
Collection #
185 
Other Collectors
 
Date Collected
2 Feb 1936 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Mason 
Locality
Twanoh State Park, Hood Canal. 
Elevation
 
Coordinates
47.37345°, -122.97428° 
Site Description
Springy area. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
78. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Marion County:
Salem.
No collection date recorded. Origin: Native.
WTU 124832; Barcode: WTU-V-015426
O. B. Johnson s.n.    
78. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31843 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
124832 
Barcode
WTU-V-015426 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris spinulosa (O.F. Muell.) Watt var. dilatata auct. non (Hoffmann) Underwood.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946. Remarks: Herbarium of The University of California.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
O. B. Johnson 
Collection #
s.n. 
Other Collectors
 
Date Collected
 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Marion 
Locality
Salem. 
Elevation
 
Coordinates
 
Site Description
 
Specimen Notes
No collection date recorded. 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
79. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, King County:
Denny Creek turnoff, U.S. 10, Snoqualmie National Forest.
47.422566°, -121.450322°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Deep coniferous woods. Abundant. Large form. Voucher: h = 41. Phenology: Fertile. Origin: Native.
WTU 213545; Barcode: WTU-V-015592
W. H. Wagner 63085    25 Jun 1963
with A. R. Kruckeberg
79. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31844 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
213545 
Barcode
WTU-V-015592 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
W. H. Wagner 
Collection #
63085 
Other Collectors
A. R. Kruckeberg 
Date Collected
25 Jun 1963 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
King 
Locality
Denny Creek turnoff, U.S. 10, Snoqualmie National Forest. 
Elevation
 
Coordinates
47.422566°, -121.450322° 
Site Description
Deep coniferous woods. 
Specimen Notes
Abundant. Large form. Voucher: h = 41. 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
80. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Marion County:
Salem.
No collection date recorded. Origin: Native.
WTU 124838; Barcode: WTU-V-015425
O. B. Johnson s.n.    
80. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31846 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
124838 
Barcode
WTU-V-015425 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946. Remarks: Herbarium of The University of California.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
O. B. Johnson 
Collection #
s.n. 
Other Collectors
 
Date Collected
 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Marion 
Locality
Salem. 
Elevation
 
Coordinates
 
Site Description
 
Specimen Notes
No collection date recorded. 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
81. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Hood River County:
Near Cascade Locks, Columbia Gorge.
Rocky talus slopes. Phenology: Spores. Origin: Native.
WTU 3333JWT; Barcode: WTU-V-015407
J. William Thompson 5146    6 Jul 1929
81. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31847 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
3333JWT 
Barcode
WTU-V-015407 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
J. William Thompson 
Collection #
5146 
Other Collectors
 
Date Collected
6 Jul 1929 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Hood River 
Locality
Near Cascade Locks, Columbia Gorge. 
Elevation
 
Coordinates
 
Site Description
Rocky talus slopes. 
Specimen Notes
 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
82. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Hood River County:
Near Cascade Locks.
Rocky talus slopes. Origin: Native.
WTU 22914; Barcode: WTU-V-015424
J. W. Thompson 5146    6 Jul 1929
82. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31849 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
22914 
Barcode
WTU-V-015424 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946. Remarks: Herbarium of The University of California.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
J. W. Thompson 
Collection #
5146 
Other Collectors
 
Date Collected
6 Jul 1929 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Hood River 
Locality
Near Cascade Locks. 
Elevation
 
Coordinates
 
Site Description
Rocky talus slopes. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
83. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Kitsap County:
Near Mountaineers' Cabin.
47.59746°, -122.73449°. Datum: WGS 84. Coordinate uncertainty: 500 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Rich moist woods near stream. Fronds gravefully curving, about 1 meter high. Phenology: Fertile. Origin: Native.
WTU 27627; Barcode: WTU-V-015591
Geo. N. Jones 4327     Aug 1932
83. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31851 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
27627 
Barcode
WTU-V-015591 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Athyrium filix-femina (L.) Roth.
2. Dryopteris dilatata auct. non (Hoffm.n) Gray.
3. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
4. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Geo. N. Jones 
Collection #
4327 
Other Collectors
 
Date Collected
Aug 1932 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Kitsap 
Locality
Near Mountaineers' Cabin. 
Elevation
 
Coordinates
47.59746°, -122.73449° 
Site Description
Rich moist woods near stream. 
Specimen Notes
Fronds gravefully curving, about 1 meter high. 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
84. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Washington County:
Roderick Falls, near Forest Grove.
Deep Ravine. Origin: Native.
WTU 3269JWT; Barcode: WTU-V-015423
J. William Thompson 4280    19 May 1928
84. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31852 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
3269JWT 
Barcode
WTU-V-015423 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946. Remarks: Herbarium of The University of California.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
J. William Thompson 
Collection #
4280 
Other Collectors
 
Date Collected
19 May 1928 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Washington 
Locality
Roderick Falls, near Forest Grove. 
Elevation
 
Coordinates
 
Site Description
Deep Ravine. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
85. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Hood River County:
3 miles from Lost Lake.
Elev. 3300 ft. 45.489349°, -121.821695°. Coordinate uncertainty: 6500 meters.
Origin: Native.
WTU 3245JWT; Barcode: WTU-V-015422
J. William Thompson 5015    23 Jul 1928
85. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31854 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
3245JWT 
Barcode
WTU-V-015422 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
J. William Thompson 
Collection #
5015 
Other Collectors
 
Date Collected
23 Jul 1928 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Hood River 
Locality
3 miles from Lost Lake. 
Elevation
3300 ft. 
Coordinates
45.489349°, -121.821695° 
Site Description
 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
86. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Oregon, Marion County:
Below Silver Creek Falls, Cascade Mountains.
Deep woods. Origin: Native.
WTU 3336JWT; Barcode: WTU-V-015421
J. William Thompson 4982    11 Jul 1928
86. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31855 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
3336JWT 
Barcode
WTU-V-015421 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946. Remarks: Herbarium of The University of California.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
J. William Thompson 
Collection #
4982 
Other Collectors
 
Date Collected
11 Jul 1928 
Verbatim Date
 
Country
U.S.A. 
State or Province
Oregon 
County
Marion 
Locality
Below Silver Creek Falls, Cascade Mountains. 
Elevation
 
Coordinates
 
Site Description
Deep woods. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
87. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Mason County:
Twanoh State Park, Hood Canal.
47.37345°, -122.97428°. Datum: WGS 84. Coordinate uncertainty: 1000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Springy area. Phenology: Vegetative. Origin: Native.
WTU 38663; Barcode: WTU-V-015590
Harold W. Smith 185    2 Feb 1936
87. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31856 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
38663 
Barcode
WTU-V-015590 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Harold W. Smith 
Collection #
185 
Other Collectors
 
Date Collected
2 Feb 1936 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Mason 
Locality
Twanoh State Park, Hood Canal. 
Elevation
 
Coordinates
47.37345°, -122.97428° 
Site Description
Springy area. 
Specimen Notes
 
Phenology
Vegetative 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
88. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Mason County:
Twanoh State Park, Hood Canal.
47.37345°, -122.97428°. Datum: WGS 84. Coordinate uncertainty: 1000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Springy area. Phenology: Fertile. Origin: Native.
WTU 38665; Barcode: WTU-V-015589
Harold W. Smith 185    2 Feb 1936
88. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31858 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
38665 
Barcode
WTU-V-015589 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Harold W. Smith 
Collection #
185 
Other Collectors
 
Date Collected
2 Feb 1936 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Mason 
Locality
Twanoh State Park, Hood Canal. 
Elevation
 
Coordinates
47.37345°, -122.97428° 
Site Description
Springy area. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
89. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Skagit County:
5 miles southwest of Anacortes.
48.461587°, -122.688629°. Datum: WGS 84. Coordinate uncertainty: 1000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Sheltered ravine in deep woods. Phenology: Fertile. Origin: Native.
WTU 45594; Barcode: WTU-V-015528
C. L. Hitchcock 3473    15 May 1938
89. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31863 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
45594 
Barcode
WTU-V-015528 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Athyrium filix-femina (L.) Roth.
2. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by W. H. Wagner, 1963.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
C. L. Hitchcock 
Collection #
3473 
Other Collectors
 
Date Collected
15 May 1938 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Skagit 
Locality
5 miles southwest of Anacortes. 
Elevation
 
Coordinates
48.461587°, -122.688629° 
Site Description
Sheltered ravine in deep woods. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
90. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Snohomish County:
Mt. Dickerman; talus above Perry Creek.
Elev. 3000 ft. 48.063294°, -121.513675°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Shaded mossy talus. Phenology: Fertile. Origin: Native.
WTU 213552; Barcode: WTU-V-015527
W. H. Wagner 63122    28 Jun 1963
with A. R. Kruckeberg
90. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31866 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
213552 
Barcode
WTU-V-015527 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
W. H. Wagner 
Collection #
63122 
Other Collectors
A. R. Kruckeberg 
Date Collected
28 Jun 1963 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Snohomish 
Locality
Mt. Dickerman; talus above Perry Creek. 
Elevation
3000 ft. 
Coordinates
48.063294°, -121.513675° 
Site Description
Shaded mossy talus. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
91. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Idaho, Shoshone County:
At head of Cirque, Lost Lake.
47.07361°, -115.95694°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Spruce fir, steep talus. Origin: Native.
WTU 271433; Barcode: WTU-V-015429
R. F. Daubenmire 45178    10 Sep 1945
91. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31868 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
271433 
Barcode
WTU-V-015429 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris austriaca (Jacq.) Woynar.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 1 2006. Sources: FNA. Remarks: WTU Herbarium. 
Type Notes
 
Collector
R. F. Daubenmire 
Collection #
45178 
Other Collectors
 
Date Collected
10 Sep 1945 
Verbatim Date
 
Country
U.S.A. 
State or Province
Idaho 
County
Shoshone 
Locality
At head of Cirque, Lost Lake. 
Elevation
 
Coordinates
47.07361°, -115.95694° 
Site Description
Spruce fir, steep talus. 
Specimen Notes
 
Phenology
 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
92. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington
Fall's Lake.
Woods. Wood fern. Phenology: Fertile. Origin: Native.
WTU 1799; Barcode: WTU-V-015526
Rigg s.n.    11 Jun 1908
92. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31869 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
1799 
Barcode
WTU-V-015526 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris spinulosa (O.F. Muell.) Watt var. dilatata auct. non (Hoffmann) Underwood.
    Annotated by T. C. Frye.
2. Dryopteris dilatata auct. non (Hoffm.n) Gray.
    Annotated by J. W. Thompson.
3. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
4. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Rigg 
Collection #
s.n. 
Other Collectors
 
Date Collected
11 Jun 1908 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
 
Locality
Fall's Lake. 
Elevation
 
Coordinates
 
Site Description
Woods. 
Specimen Notes
Wood fern. 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
93. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, San Juan County:
Mt. Constitution, Orcas Island.
48.672778°, -122.841111°. Datum: WGS 84. Coordinate uncertainty: 3000 meters. Georeferenced by WTU Staff. Coordinate Source: Georeferenced.
Phenology: Fertile. Origin: Native.
WTU 34857; Barcode: WTU-V-015525
T. C. Frye s.n.    19 Jul 1904
93. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31873 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
34857 
Barcode
WTU-V-015525 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
T. C. Frye 
Collection #
s.n. 
Other Collectors
 
Date Collected
19 Jul 1904 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
San Juan 
Locality
Mt. Constitution, Orcas Island. 
Elevation
 
Coordinates
48.672778°, -122.841111° 
Site Description
 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
94. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Jefferson County:
Olympic Forest. East Fork Quinault Near Fire Creek.
Elev. 600 ft. 47.851165°, -124.253724°. Datum: WGS 84. Coordinate uncertainty: 5000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Flat. Soil: Shallow clay loam. Type: Timber, Browse. Density: 3. Associated: Lady Fern, Oxalis. Moist woods. Not common. 8" - 16". Phenology: Fertile. Origin: Native.
WTU 322798; Barcode: WTU-V-015509
John E. Schwartz 26 A    20 Aug 1935
94. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31875 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
322798 
Barcode
WTU-V-015509 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
John E. Schwartz 
Collection #
26 A 
Other Collectors
 
Date Collected
20 Aug 1935 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Jefferson 
Locality
Olympic Forest. East Fork Quinault Near Fire Creek. 
Elevation
600 ft. 
Coordinates
47.851165°, -124.253724° 
Site Description
Flat. Soil: Shallow clay loam. Type: Timber, Browse. Density: 3. Associated: Lady Fern, Oxalis. Moist woods. 
Specimen Notes
Not common. 8" - 16". 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
95. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Whatcom County:
Base of Panorama Point, near Mt. Baker Lodge.
Elev. 4200 ft. 48.85194°, -121.68389°. Georeferenced by Ed Alverson. Coordinate Source: Inferred from duplicate specimen.
Phenology: Fertile. Origin: Native.
WTU 3198JWT; Barcode: WTU-V-015510
J. William Thompson 5720 1/2    13 Aug 1930
95. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31880 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
3198JWT 
Barcode
WTU-V-015510 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
J. William Thompson 
Collection #
5720 1/2 
Other Collectors
 
Date Collected
13 Aug 1930 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Whatcom 
Locality
Base of Panorama Point, near Mt. Baker Lodge. 
Elevation
4200 ft. 
Coordinates
48.85194°, -121.68389° 
Site Description
 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
96. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, San Juan County:
Mount Constitution, Mountain Lake, Orcas Island.
48.672778°, -122.841111°. Datum: WGS 84. Coordinate uncertainty: 3000 meters. Georeferenced by WTU Staff. Coordinate Source: Georeferenced.
Phenology: Fertile. Origin: Native.
WTU 124833; Barcode: WTU-V-015511
T. C. Frye s.n.    19 Jul 1904
96. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31883 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
124833 
Barcode
WTU-V-015511 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
T. C. Frye 
Collection #
s.n. 
Other Collectors
 
Date Collected
19 Jul 1904 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
San Juan 
Locality
Mount Constitution, Mountain Lake, Orcas Island. 
Elevation
 
Coordinates
48.672778°, -122.841111° 
Site Description
 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
97. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Snohomish County:
Cascade Mountains. Along Perry Creek.
Elev. 3500 ft. 48.063294°, -121.513675°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Along wooded creek. Phenology: Fertile. Origin: Native.
WTU 173349; Barcode: WTU-V-015512
J. W. Thompson 17484    30 Jun 1953
97. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31884 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
173349 
Barcode
WTU-V-015512 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
J. W. Thompson 
Collection #
17484 
Other Collectors
 
Date Collected
30 Jun 1953 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Snohomish 
Locality
Cascade Mountains. Along Perry Creek. 
Elevation
3500 ft. 
Coordinates
48.063294°, -121.513675° 
Site Description
Along wooded creek. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
98. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Pierce County:
Ranger Camp, Mount Rainier.
Elev. 4500 ft.
Phenology: Fertile. Origin: Native.
WTU 22145; Barcode: WTU-V-015513
J. M. Grant s.n.     Aug 1925
98. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31886 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
22145 
Barcode
WTU-V-015513 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris spinulosa (O.F. Muell.) Kunze.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.  
Type Notes
 
Collector
J. M. Grant 
Collection #
s.n. 
Other Collectors
 
Date Collected
Aug 1925 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Pierce 
Locality
Ranger Camp, Mount Rainier. 
Elevation
4500 ft. 
Coordinates
 
Site Description
 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
99. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Pierce County:
Trail to Spray Park, Mount Rainier.
Elev. 4000 ft. 46.91556°, -121.82444°. Datum: WGS 84. Coordinate uncertainty: 10000 meters. Georeferenced by WTU Staff. Coordinate Source: GeoLocate.
Deep woods. Phenology: Fertile. Origin: Native.
WTU 3200JWT; Barcode: WTU-V-015502
J. William Thompson 5463    27 Jul 1930
99. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31889 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
3200JWT 
Barcode
WTU-V-015502 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Dryopteris dilatata auct. non (Hoffm.n) Gray.
2. Dryopteris austriaca (Jacq.) Woynar.
    Annotated by Carlotta C. Hall, 1946.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
J. William Thompson 
Collection #
5463 
Other Collectors
 
Date Collected
27 Jul 1930 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Pierce 
Locality
Trail to Spray Park, Mount Rainier. 
Elevation
4000 ft. 
Coordinates
46.91556°, -121.82444° 
Site Description
Deep woods. 
Specimen Notes
 
Phenology
Fertile 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

Dryopteridaceae
100. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
U.S.A., Washington, Pierce County:
Mount Rainier.
46.852827°, -121.760441°. Datum: WGS 84. Coordinate uncertainty: 17500 meters. Georeferenced by Joseph Stormer. Coordinate Source: GeoLocate.
Phenology: Spores. Origin: Native.
WTU 90307; Barcode: WTU-V-015514
Emma A. Shumway s.n.     Aug 1889
100. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy
Database ID
31891 
Herbarium
WTU 
Collection
Vascular Plants 
Dataset
 
Accession
90307 
Barcode
WTU-V-015514 
Other Numbers
 
Family
Dryopteridaceae 
Name on Specimen
Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy 
Accepted Name
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy 
Annotation History
1. Aspidium spinulosum var. dilatatum Sw.
2. Dryopteris austriaca (Jacq.) Woynar.
3. Dryopteris expansa (C. Presl) Fraser-Jenkins & Jermy.
    Annotated by Ed Alverson, 12 2004.  
Type Notes
 
Collector
Emma A. Shumway 
Collection #
s.n. 
Other Collectors
 
Date Collected
Aug 1889 
Verbatim Date
 
Country
U.S.A. 
State or Province
Washington 
County
Pierce 
Locality
Mount Rainier. 
Elevation
 
Coordinates
46.852827°, -121.760441° 
Site Description
 
Specimen Notes
 
Phenology
Spores 
Origin
Native 
Cultivated?
No 
Imaged?
Yes 

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Database Field Descriptions:
Database ID
Each of our backend databases assigns a unique and persistent integer ID to each specimen record. A GUID (globally unique identifier) can be formed from this by combining it with the herbarium and collection acronyms (example: WTU-VP-12733).
Herbarium
Every herbarium is referred to by a unique acronym. Ours is WTU, which stands for Washington Territorial University. Our database also includes some specimens housed at NOCA (North Cascades National Park Herbarium) and MORA (Mount Rainier National Park Herbarium). For a complete listing of herbarium acronyms, see Index Herbariorum.
Collection
WTU's specimens are physically housed in separate collections according to taxonomic groups: Vascular Plants, Bryophytes, Algae, Lichens, and Fungi.
Dataset
Some specimens are grouped into "Datasets," representing such things as incorporated herbaria or major collecting projects.
Accession
Each of WTU's Vascular Plant specimens is assigned an accession number. Our other collections lack accession numbers (see Barcode instead). Ideally, each specimen would have a unique accession number, but this fails in practice. Some accession numbers include "JWT" after the number; these indicate specimens originating from the herbarium of J. William Thompson that were integrated into WTU's collections but not re-accessioned with WTU numbers. Accession numbers followed by an "A", "B", or other single letter represent sheets with multiple collections mounted on them, all using the same accession number.
Barcode
Our Bryophyte, Algae, Lichen, Fungi, and some Vascular Plant specimens are assigned pre-printed barcode stickers. These follow the format "WTU-C-######", where C represents a code for the collection (one of: V, B, A, L, F) and ###### represents a zero-padded, six-digit number.
Other Numbers
Specimens collected from North Cascades and Mount Rainier National Parks are assigned National Park Service accession and catalog numbers. These numbers are listed here.
Family
The scientific name of the family. Our database generally uses AGPIII and other current sources for vascular plant family names. However, our physical Vascular Plant collection is organized by the older Englerian sequence.
Accepted Name
For Vascular Plants, this gives the accepted name by which the specimen is currently known. The accepted name is derived by looking up the most recent annotation name in a synonymy table and finding its corresponding accepted name.
Annotation History
The sequence of annotations and scientific names given on the physical specimen is listed here, in order from the first to most recent. We treat the name on the original label as the first annotation.
Type Notes
If this is a type specimen, then information about its type status (holotype, isotype, etc.) and the name(s) for which it is a type is provided here.
Collector
The person who collected this specimen, as given on the label, and whose name is associated with the collection number on the label. If no collector name is given, we enter "unknown" into this field.
Collection #
A unique number assigned to the specimen by the collector. Different collectors use different numbering schemes; these may include the addition of prefixes (such as their initials or the year) or suffixes (such as an A or B). In the database, any prefixes and suffixes are included in the Collector's Number field exactly as they appear on the specimen label. When no collector number is given, we type "s.n." in this field.
Other Collectors
Other collectors that are listed on the label along with the primary collector, but whose collection number was not assigned to this specimen.
Date Collected
The date on which the specimen was collected, split up between day, month, and year in the database. Months are stored as nubmers (1-12) but displayed as three-letter abbreviations. Years are alway 4-digits long. Occasionally the label may give a range of dates. In such cases, the first date in the range is used as the collection date, and the entire collection date is entered into the Verbatim Date field as it appears on the label. If no date is given, then the date fields are left blank and a note is made in the Verbatim Date field.
Verbatim Date
The literal date string as displayed on the label; only used for dates that cannot be represented by a single day, month, and year.
Country
Name of the country from which the specimen was collected. Specimens from the United States of America are entered as "U.S.A."
State or Province
Name of the state or province from which the specimen was collected, if given on the label. U.S. states are spelled out in full.
County
Name of the county, municipality, borough, parish, or equivalent political unit from which the specimen was collected, if given on the label. The designation (such as "county") is not included in the field value.
Locality
A text description of the location where the specimen was collected, as given on the specimen label. May include national park, national forest, and other land use designations. Older collections often have short and imprecise locality data. Detailed locality data for rare species (threatened, endangered, sensitive, watch, ...) is included in our database but omitted from the search results.
Elevation
Elevations may be given either in feet or meters, and may include a minimum and maximum elevation, representing a range.
Coordinates
May include TRS/PLSS (township, range, section, quarter, etc.), latitude/longitude, and UTM. Coordinates for arare species are included in our database but omittedfrom the search results.
Site Description
This field includes all label information characterizing the site, such habitat, vegetation zone/association, substrate, slope, moisture, associated species, and so on - as the information appears on the specimen label.
Specimen Notes
Includes any characteristics pertaining to the plant specimen itself, such as flower color, fruit length, and plant height, along with information about the species, such as abundance and the precise location and micro-habitat where collected. Also includes any notes or text on the specimen that does not better fit into another field.
Phenology
Specifies the sexual maturity/condition of the specimen, such as flowering, fruiting, cone-bearing, spore-bearing, fertile, or sterile. Phenology is generally not given on the label; instead the person entering the record examines the specimen and selects the appropriate value.
Origin
Specifies whether the specimen is native or introduced to the state or country where it was collected. Origin data is rarely found on the specimen label; instead, it is entered during data entry either from a lookup table or from the date entry personnel's own knowledge. Origin data should be considered supplemental information, and should not be relied on as authoritative.
Cultivated
A Yes/No flag indicating if this is a cultivated specimen. Cultivated specimens generally do not represent valid distribution records and should be excluded from range maps or other geographic analyses.


Disclaimer:

WTU makes a reasonable effort to prevent and correct data entry errors; however, we cannot guarantee that our database is free from typos, misidentifications, or inaccurate label data. Please use the information at your own discretion. Error corrections and comments can be sent to Ben Legler or David Giblin.